By Paul Haas
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Extra resources for an introduction to the chemistry of plant products
A general account of chlorophyll, its chemistry and constitution, will be found in Vol. I. § Irving : " Ann. ," 1910, 34,, 805. CHLOROPHYLL 43 I per cent, of the activity subsequently developed. Carbon assimilation thus begins only when the leaves are fully green and develops very quickly ; wherefore it follows that the first origin of this function is not correlative to the amount of chlorophyll produced, or, in other words, that the amount of chlorophyll is not a conditioning factor in the early stages of carbon assimilation.
Comparable observations \ave been made by Yabusoe * and also by Lundegardh f who found the temperature coefficient to be 1-03-1-45 a t 3°c-35° in "a strong light and with an atmosphere enriched to 1-2 per cent, of carbon dioxide. The maximum rate occurred at 350 C. and provided that the initial heating was not too prolonged, the time factor was ill-defined. Blackman observed that under strong illumination or in a high concentration of carbon dioxide, a rise of io° in temperature, from 15° to 250, roughly doubles the velocity of photosynthesis ; this is termed by Warburg % the "Blackman Reaction " and it indicates that in the conditions specified, the rate of carbon assimilation is determined by a chemical rather than a photochemical reaction, for, as has been seen, with a low light intensity a rise in temperature does not affect the rate of the reaction.
F See Palladia: **Ber. deut. bat. Cka," 1891, ip» 229. Mansky: "Itiodteoi. " 1922,132, x8. **Se®Ewart; "Ajia. Bot,/* 1S97, u r 459; 1S9S, 1:2, 379. Pantaaoili: "J&kr. wisa. Bat," 1903, 3^, 167. Urspraiig: *f Ber. d«nt. bot, & " M* 57- INFLUENCE OF ILLUMINATION 31 be lost by reflection, absorption or transmission. Of the energy absorbed by the leaf, many have shown that a small proportion only, and this in varying quantity, is used in carbon assimilation. Owing to ignorance of certain factors and the degree of their significance in the sequence of carbon assimilation, it is not possible to give a satisfactory account of the energy relationships of the plant.